1.10 Overwatch, 2nd para says
Quote:
A formation that is on overwatch may choose to shoot immediately after an enemy formation completes a move or unloads troops, and before the target either shoots or assaults.
There is nothing preventing units that are now in B-B from shooting, so all defending units that have LoS and range to an attacking unit may fire, even if that unit is now in B-B with a defender. So, basically everyone can shoot under the usual restrictions.
One of the more typical restrictions is terrain which may limit or prohibit some units from firing. Rarely, some or all of the attackers my now be outside the arc of fire (if attacking some WE or titans that were on OW).
Note RAW, shooting is done at the end of the engage move, preventing shooting at units that are now 'hidden' even if they could be seen at the start of their movement, although some people play a house rule that moving through an arc of fire makes a unit eligible for shooting.